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Faculty and Staff
ZHOU Zhekun
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Mailing Address:
XTBG, Menglun, Mengla, Yunnan, 666303,China


Major research interests include 1) paleoenvironment and biodiversity change caused by major geological events such as the uplift of the Himalayas and the onset of the Asian monsoon system, and its corresponding effect on the distribution of plants; 2) morphological responses of plants to climate change and consequent changes in plant distribution; 3) reconstruction of the paleo-CO2 environment, and paleoelevation; 4). taxonomy, fossil history, modern distribution, phylogeny, conservation, and ecology of the Fagaceae and other main elements of eastern Aisa



1.         Zhou Z, Hu JJ, Wen J, Sun H: Morphometric, phylogenetic and biogeographic analyses of Pyrularia (Santalales), a parasitic disjunct lineage between eastern Asia and eastern North America. Taxon 2019, 68(1):47-71. 

2.         Zhao F, Fan ZX, Su T, Li SF, Tang H, Spicer TEV, Zhou ZK: Tree-ring delta O-18 inferred spring drought variability over the past 200 years in the Hengduan Mountains, Southwest China. Palaeogeography Palaeoclimatology Palaeoecology 2019, 518:22-33. 

3.         Xu H, Su T, Zhou ZK: Leaf and infructescence fossils of Alnus (Betulaceae) from the late Eocene of the southeastern Qinghai-Tibetan Plateau. Journal of Systematics and Evolution 2019, 57(2):105-113. 

4.         Xu CL, Su T, Huang J, Huang YJ, Li SF, Zhao YS, Zhou ZK: Occurrence of Christella (Thelypteridaceae) in Southwest China and its indications of the paleoenvironment of the Qinghai-Tibetan Plateau and adjacent areas. Journal of Systematics and Evolution 2019, 57(2):169-179. 

5.         Wang L, Kunzmann L, Su T, Xing YW, Zhang ST, Wang YQ, Zhou ZK: The disappearance of Metasequoia (Cupressaceae) after the middle Miocene in Yunnan, Southwest China: Evidences for evolutionary stasis and intensification of the Asian monsoon. Review of Palaeobotany and Palynology 2019, 264:64-74. 

6.         Su T, Spicer RA, Li SH, Xu H, Huang J, Sherlock S, Huang YJ, Li SF, Wang L, Jia LB et al: Uplift, climate and biotic changes at the Eocene-Oligocene transition in south-eastern Tibet. National Science Review 2019, 6(3):495-504. 

7.         Su T, Farnsworth A, Spicer RA, Huang J, Wus FX, Liu J, Li SF, Xing YW, Huang YJ, Deng WYD et al: No high Tibetan Plateau until the Neogene. Science Advances 2019, 5(3). 

8.         Liu J, Su T, Spicer RA, Tang H, Deng WYD, Wu FX, Srivastava G, Spicer T, Do VT, Deng T et al: Biotic interchange through lowlands of Tibetan Plateau suture zones during Paleogene. Palaeogeography Palaeoclimatology Palaeoecology 2019, 524:33-40. 

9.         Jia LB, Su T, Huang YJ, Wu FX, Deng T, Zhou ZK: First fossil record of Cedrelospermum (Ulmaceae) from the Qinghai-Tibetan Plateau: Implications for morphological evolution and biogeography. Journal of Systematics and Evolution 2019, 57(2):94-104. 

10.       Huang YJ, Zhu H, Momohara A, Jia LB, Zhou ZK: Fruit fossils of Rosoideae (Rosaceae) from the late Pliocene of northwestern Yunnan, Southwest China. Journal of Systematics and Evolution 2019, 57(2):180-189. 

11.       Chen WY, Su T, Jia LB, Zhou ZK: The relationship between leaf physiognomy and climate based on a large modern dataset: Implications for palaeoclimate reconstructions in China. Palaeogeography Palaeoclimatology Palaeoecology 2019, 527:1-13. 

12.       Zhou ZK: A tribute to my supervisor Professor Zhengyi Wu. Protein & Cell 2018, 9(10):827-831. 

13.       Zhang QY, Huang J, Jia LB, Su T, Zhou ZK, Xing YW: Miocene Ulmus fossil fruits from Southwest China and their evolutionary and biogeographic implications. Review of Palaeobotany and Palynology 2018, 259:198-206. 

14.       Su T, Li SF, Tang H, Huang YJ, Li SH, Deng CL, Zhou ZK: Hemitrapa Miki (Lythraceae) from the earliest Oligocene of southeastern Qinghai-Tibetan Plateau and its phytogeographic implications. Review of Palaeobotany and Palynology 2018, 257:57-63. 

15.       Panthi S, Brauning A, Zhou ZK, Fan ZX: Growth response of Abies georgei to climate increases with elevation in the central Hengduan Mountains, southwestern China. Dendrochronologia 2018, 47:1-9. 

16.       Linnemann U, Su T, Kunzmann L, Spicer RA, Ding WN, Spicer TEV, Zieger J, Hofmann M, Moraweck K, Gartner A et al: New U-Pb dates show a Paleogene origin for the modern Asian biodiversity hot spots. Geology 2018, 46(1):3-6. 

17.       Liang XQ, Lu P, Zhang JW, Su T, Zhou ZK: First fossils of Zygogynum from the Middle Miocene of Central Yunnan, Southwest China, and their palaeobiogeographic significance. Palaeoworld 2018, 27(3):399-409. 

18.       Li SF, Xing YW, Valdes PJ, Huang YJ, Su T, Farnsworth A, Lunt DJ, Tang H, Kennedy AT, Zhou ZK: Oligocene climate signals and forcings in Eurasia revealed by plant macrofossil and modelling results. Gondwana Research 2018, 61:115-127. 

19.       Huang J, Su T, Jia LB, Spicer T, Zhou ZK: A fossil fig from the Miocene of southwestern China: Indication of persistent deep time karst vegetation. Review of Palaeobotany and Palynology 2018, 258:133-145. 

20.       Huang HS, Su T, Zhou ZK: Fossil leaves of Buxus (Buxaceae) from the Upper Pliocene of Yunnan, SW China. Palaeoworld 2018, 27(2):271-281. 

21.       Engel MS, Wang B, Alqarni AS, Jia LB, Su T, Zhou ZK, Wappler T: A primitive honey bee from the Middle Miocene deposits of southeastern Yunnan, China (Hymenoptera, Apidae). Zookeys 2018(775):117-129. 

22.       Ding WN, Kunzmann L, Su T, Huang J, Zhou ZK: A new fossil species of Cryptomeria (Cupressaceae) from the Rupelian of the Luhe Basin, Yunnan, East Asia: Implications for palaeobiogeography and palaeoecology. Review of Palaeobotany and Palynology 2018, 248:41-51. 

23.       Ding WN, Huang J, Su T, Xing YW, Zhou ZK: An early Oligocene occurrence of the palaeoendemic genus Dipteronia (Sapindaceae) from Southwest China. Review of Palaeobotany and Palynology 2018, 249:16-23. 

24.       Xu CL, Huang J, Su T, Zhang XC, Li SF, Zhou ZK: The first megafossil record of Goniophlebium (Polypodiaceae) from the Middle Miocene of Asia and its paleoecological implications. Palaeoworld 2017, 26(3):543-552. 

25.       Tiwari A, Fan ZX, Jump AS, Zhou ZK: Warming induced growth decline of Himalayan birch at its lower range edge in a semi-arid region of Trans-Himalaya, central Nepal. Plant Ecology 2017, 218(5):621-633. 

26.       Tiwari A, Fan ZX, Jump AS, Li SF, Zhou ZK: Gradual expansion of moisture sensitive Abies spectabilis forest in the Trans-Himalayan zone of central Nepal associated with climate change. Dendrochronologia 2017, 41:34-43. 

27.       Panthi S, Brauning A, Zhou ZK, Fan ZX: Tree rings reveal recent intensified spring drought in the central Himalaya, Nepal. Global and Planetary Change 2017, 157:26-34. 

28.       Meng HH, Su T, Gao XY, Li J, Jiang XL, Sun H, Zhou ZK: Warm-cold colonization: response of oaks to uplift of the Himalaya-Hengduan Mountains. Molecular Ecology 2017, 26(12):3276-3294. 

29.       Liang XQ, Lu P, Tiwari A, Su T, Zhou ZK: New fossil record of Cladium (Cyperaceae) from the Middle Miocene of Zhenyuan, SW China, and the palaeobiogeographical history of the genus. Review of Palaeobotany and Palynology 2017, 237:1-9. 

30.       Li SF, Hughes AC, Su T, Anberree JL, Oskolski AA, Sun M, Ferguson DK, Zhou ZK: Fire dynamics under monsoonal climate in Yunnan, SW China: past, present and future. Palaeogeography Palaeoclimatology Palaeoecology 2017, 465:168-176. 

31.       Kang HB, Oskolski AA, Jacques FMB, Wang YH, Zhou ZK: Lignified woods of Pinus (Pinaceae) from the late Miocene of central Yunnan, China, and their biogeographic and paleoclimatic implications. Palaeoworld 2017, 26(3):553-565. 

32.       Jia LB, Huang YJ, Sun H, Su T, Huang J, Zhou ZK: First fossil of Pterolobium (Leguminosae) from the Middle Miocene Yunnan, South China. Review of Palaeobotany and Palynology 2017, 242:21-32. 

33.       Huang YJ, Ji XP, Su T, Deng CL, Ferguson DK, Yu TS, Yang X, Sun H, Zhou ZK: Habitat, climate and potential plant food resources for the late Miocene Shuitangba hominoid in Southwest China: Insights from carpological remains. Palaeogeography Palaeoclimatology Palaeoecology 2017, 470:63-71. 

34.       Huang J, Shi GL, Su T, Zhou ZK: Miocene Exbucklandia (Hamamelidaceae) from Yunnan, China and its biogeographic and palaeoecologic implications. Review of Palaeobotany and Palynology 2017, 244:96-106. 

35.       Zhu H, Huang YJ, Ji XP, Su T, Zhou ZK: Continuous existence of Zanthoxylum (Rutaceae) in Southwest China since the Miocene. Quaternary International 2016, 392:224-232. 

36.       Xu H, Su T, Zhang ST, Deng M, Zhou ZK: The first fossil record of ring-cupped oak (Quercus L. subgenus Cyclobalanopsis (Oersted) Schneider) in Tibet and its paleoenvironmental implications. Palaeogeography Palaeoclimatology Palaeoecology 2016, 442:61-71. 

37.       Xia K, Tan HY, Turkington R, Hu JJ, Zhou ZK: Desiccation and post-dispersal infestation of acorns of Quercus schottkyana, a dominant evergreen oak in SW China. Plant Ecology 2016, 217(11):1369-1378. 

38.       Xia K, Harrower WL, Turkington R, Tan HY, Zhou ZK: Pre-dispersal strategies by Quercus schottkyana to mitigate the effects of weevil infestation of acorns. Scientific Reports 2016, 6. 

39.       Sun M, Su T, Zhang SB, Li SF, Anberree-Lebreton J, Zhou ZK: Variations in Leaf Morphological Traits of Quercus guyavifolia (Fagaceae) were Mainly Influenced by Water and Ultraviolet Irradiation at High Elevations on the Qinghai-Tibet Plateau, China. International Journal of Agriculture and Biology 2016, 18(2):266-273. 

40.       Su T, Huang YJ, Meng J, Zhang ST, Huang J, Zhou ZK: A Miocene leaf fossil record of Rosa (R-fortuita n. sp.) from its modern diversity center in SW China. Palaeoworld 2016, 25(1):104-115. 

41.       Lu J, Turkington R, Zhou ZK: The effects of litter quantity and quality on soil nutrients and litter invertebrates in the understory of two forests in southern China. Plant Ecology 2016, 217(11):1415-1428. 

42.       Liang XQ, Ferguson DK, Su T, Zhou ZK: Fossil leaves of Populus from the Middle Miocene of Yunnan, SW China. Journal of Systematics and Evolution 2016, 54(3):264-271. 

43.       Liang XQ, Ferguson DK, Jacques FMB, Su T, Wang L, Zhou ZK: A new Celastrus species from the middle Miocene of Yunnan, China and its palaeoclimatic and palaeobiogeographic implications. Review of Palaeobotany and Palynology 2016, 225:43-52. 

44.       Li SF, Jacques FMB, Spicer RA, Su T, Spicer TEV, Yang J, Zhou ZK: Artificial neural networks reveal a high-resolution climatic signal in leaf physiognomy. Palaeogeography Palaeoclimatology Palaeoecology 2016, 442:1-11. 

45.       Lebreton-Anberree J, Li SH, Li SF, Spicer RA, Zhang ST, Su T, Deng CL, Zhou ZK: Lake geochemistry reveals marked environmental change in Southwest China during the Mid Miocene Climatic Optimum. Science Bulletin 2016, 61(11):897-910. 

46.       Huang J, Su T, Lebereton-Anberree J, Zhang ST, Zhou ZK: The oldest Mahonia (Berberidaceae) fossil from East Asia and its biogeographic implications. Journal of Plant Research 2016, 129(2):209-223. 

47.       Hu JJ, Luo CC, Turkington R, Zhou ZK: Effects of herbivores and litter on Lithocarpus hancei seed germination and seedling survival in the understorey of a high diversity forest in SW China. Plant Ecology 2016, 217(11):1429-1440. 

48.       Zhu H, Jacques FMB, Wang L, Xiao XH, Huang YJ, Zhou ZK: Fossil endocarps of Aralia (Araliaceae) from the upper Pliocene of Yunnan in southwest China, and their biogeographical implications. Review of Palaeobotany and Palynology 2015, 223:94-103. 

49.       Zhou ZK, Jiang Y, Wang Z, Gou ZH, Lyu J, Li WY, Yu YJ, Shu LP, Zhao YJ, Ma YM et al: Resequencing 302 wild and cultivated accessions identifies genes related to domestication and improvement in soybean. Nature Biotechnology 2015, 33(4):408-U125. 

50.       Zhang JW, D'Rozario A, Adams JM, Liang XQ, Jacques FMB, Su T, Zhou ZK: The occurrence of Pinus massoniana Lambert (Pinaceae) from the upper Miocene of Yunnan, SW China and its implications for paleogeography and paleoclimate. Review of Palaeobotany and Palynology 2015, 215:57-67. 


52.       Yang J, Spicer RA, Spicer TEV, Arens NC, Jacques FMB, Su T, Kennedy EM, Herman AB, Steart DC, Srivastava G et al: Leaf form-climate relationships on the global stage: an ensemble of characters. Global Ecology and Biogeography 2015, 24(10):1113-1125. 

53.       Xu QQ, Qiu J, Jin JH, Zhou ZK: Eocene Podocarpium (Leguminosae) from South China and its biogeographic implications (vol 6, 938, 2015). Frontiers in Plant Science 2015, 6. 

54.       Xia K, Daws MI, Zhou ZK, Pritchard HW: Habitat-linked temperature requirements for fruit germination in Quercus species: A comparative study of Quercus subgenus Cyclobalanopsis (Asian evergreen oaks) and Quercus subgenus Quercus. South African Journal of Botany 2015, 100:108-113. 

55.       Wang YQ, Momohara A, Wang L, Lebreton-Anberree J, Zhou ZK: Evolutionary History of Atmospheric CO2 during the Late Cenozoic from Fossilized Metasequoia Needles. Plos One 2015, 10(7). 

56.       Wang HB, Bondarenko OV, Jacques FMB, Wang YH, Zhou ZK: A New Species of Tsuga (Pinaceae) based on Lignified Wood from the Late Miocene of Central Yunnan, China, and Its Paleoenvironmental Implications. Acta Geologica Sinica-English Edition 2015, 89(5):1429-1439. 

57.       Su T, Wilf P, Huang YJ, Zhang ST, Zhou ZK: Peaches Preceded Humans: Fossil Evidence from SW China. Scientific Reports 2015, 5. 

58.       Su T, Adams JM, Wappler T, Huang YJ, Jacques FMB, Liu YS, Zhou ZK: Resilience of plant-insect interactions in an oak lineage through Quaternary climate change. Paleobiology 2015, 41(1):174-186. 

59.       Meng HH, Su T, Huang YJ, Zhu H, Zhou ZK: Late Miocene Palaeocarya (Engelhardieae: Juglandaceae) from Southwest China and its biogeographic implications. Journal of Systematics and Evolution 2015, 53(6):499-511. 

60.       Li YJ, Oskolski AA, Jacques FMB, Zhou ZK: NEW MIDDLE MIOCENE FOSSIL WOOD OF WATARIA (MALVACEAE) FROM SOUTHWEST CHINA. Iawa Journal 2015, 36(3):345-357. 

61.       Li SF, Mao LM, Spicer RA, Lebreton-Anberree J, Su T, Sun M, Zhou ZK: Late Miocene vegetation dynamics under monsoonal climate in southwestern China. Palaeogeography Palaeoclimatology Palaeoecology 2015, 425:14-40. 

62.       Jia LB, Manchester SR, Su T, Xing YW, Chen WY, Huang YJ, Zhou ZK: First occurrence of Cedrelospermum (Ulmaceae) in Asia and its biogeographic implications. Journal of Plant Research 2015, 128(5):747-761. 

63.       Jacques FMB, Shi GL, Su T, Zhou ZK: A tropical forest of the middle Miocene of Fujian (SE China) reveals Sino-Indian biogeographic affinities. Review of Palaeobotany and Palynology 2015, 216:76-91. 

64.       Huang YJ, Ji XP, Su T, Wang L, Deng CL, Li WQ, Luo HF, Zhou ZK: Fossil seeds of Euryale (Nymphaeaceae) indicate a lake or swamp environment in the late Miocene Zhaotong Basin of southwestern China. Science Bulletin 2015, 60(20):1768-1777. 

65.       Huang YJ, Jacques FMB, Su T, Ferguson DK, Tang H, Chen WY, Zhou ZK: Distribution of Cenozoic plant relicts in China explained by drought in dry season. Scientific Reports 2015, 5. 

66.       Huang YJ, Jacques FMB, Liu YSC, Su T, Ferguson DK, Xing YW, Zhou ZK: Rubus (Rosaceae) diversity in the late Pliocene of Yunnan, southwestern China. Geobios 2015, 48(6):439-448. 

67.       Huang YJ, Chen WY, Jacques FMB, Liu YSC, Utescher T, Su T, Ferguson DK, Zhou ZK: Late Pliocene temperatures and their spatial variation at the southeastern border of the Qinghai-Tibet Plateau. Journal of Asian Earth Sciences 2015, 111:44-53. 

68.       Hu JJ, Xing YW, Turkington R, Jacques FMB, Su T, Huang YJ, Zhou ZK: A new positive relationship between pCO(2) and stomatal frequency in Quercus guyavifolia (Fagaceae): a potential proxy for palaeo-CO2 levels. Annals of Botany 2015, 115(5):777-788. 


70.       Zhang MG, Zhou ZK, Chen WY, Cannon CH, Raes N, Slik JWF: Major declines of woody plant species ranges under climate change in Yunnan, China. Diversity and Distributions 2014, 20(4):405-415. 

71.       Xiang XG, Wang W, Li RQ, Lin L, Liu Y, Zhou ZK, Li ZY, Chen ZD: Large-scale phylogenetic analyses reveal fagalean diversification promoted by the interplay of diaspores and environments in the Paleogene. Perspectives in Plant Ecology Evolution and Systematics 2014, 16(3):101-110. 


73.       Meng HH, Jacques FMB, Su T, Huang YJ, Zhang ST, Ma HJ, Zhou ZK: New Biogeographic insight into Bauhinia s.l. (Leguminosae): integration from fossil records and molecular analyses. Bmc Evolutionary Biology 2014, 14. 

74.       Jacques FMB, Su T, Spicer RA, Xing YW, Huang YJ, Zhou ZK: Late Miocene southwestern Chinese floristic diversity shaped by the southeastern uplift of the Tibetan Plateau. Palaeogeography Palaeoclimatology Palaeoecology 2014, 411:208-215. 

75.       Hu Q, Xing YW, Hu JJ, Huang YJ, Ma HJ, Zhou ZK: Evolution of stomatal and trichome density of the Quercus delavayi complex since the late Miocene. Chinese Science Bulletin 2014, 59(3):310-319. 

76.       Chen WY, Su T, Adams JM, Jacques FMB, Ferguson DK, Zhou ZK: Large-scale dataset from China gives new insights into leaf '>Palaeogeography Palaeoclimatology Palaeoecology 2014, 402:73-80. 

77.       Xing YW, Liu YSC, Su T, Jacques FMB, Zhou ZK: A new Tsuga species from the upper Miocene of Yunnan, southwestern China and its palaeogeographic significance. Palaeoworld 2013, 22(3-4):159-167. 

78.       Xing YW, Hu JJ, Jacques FMB, Wang L, Su T, Huang YJ, Liu YS, Zhou ZK: A new Quercus species from the upper Miocene of southwestern China and its ecological significance. Review of Palaeobotany and Palynology 2013, 193:99-109. 

79.       Wang WM, Zhou ZK: Neogene climate evolution in Eastern Eurasia with special reference to floral changes under the influence of monsoon. Palaeoworld 2013, 22(3-4):73-76. 

80.       Wang L, Jacques FMB, Su T, Xing YW, Zhang ST, Zhou ZK: The earliest fossil bamboos of China (middle Miocene, Yunnan) and their biogeographical importance. Review of Palaeobotany and Palynology 2013, 197:253-265. 

81.       Su T, Spicer RA, Liu YS, Huang YJ, Xing YW, Jacques FMB, Chen WY, Zhou ZK: Regional constraints on leaf physiognomy and precipitation regression models: a case study from China. Bulletin of Geosciences 2013, 88(3):595-608. 

82.       Su T, Liu YS, Jacques FMB, Huang YJ, Xing YW, Zhou ZK: The intensification of the East Asian winter monsoon contributed to the disappearance of Cedrus (Pinaceae) in southwestern China. Quaternary Research 2013, 80(2):316-325. 

83.       Su T, Jacques FMB, Spicer RA, Liu YS, Huang YJ, Xing YW, Zhou ZK: Post-Pliocene establishment of the present monsoonal climate in SW China: evidence from the late Pliocene Longmen megaflora. Climate of the Past 2013, 9(4):1911-1920. 

84.       Su T, Jacques FMB, Ma HJ, Zhou ZK: Fossil fruits of Ailanthus confucii from the Upper Miocene of Wenshan, Yunnan Province, southwestern China. Palaeoworld 2013, 22(3-4):153-158. 

85.       Jacques FMB, Su T, Zhou ZK: The first fossil Microsoroid fern (Palaeosorum ellipticum gen. et sp nov.) from the middle Miocene of Yunnan, SW China. Journal of Systematics and Evolution 2013, 51(6):758-764. 

86.       Jacques FMB, Su T, Huang YJ, Wang L, Zhou ZK: A global-scale test for monsoon indices used in palaeoclimatic reconstruction. Palaeoworld 2013, 22(3-4):93-100. 

87.       Huang YJ, Liu YS, Jacques FMB, Su T, Xing YW, Zhou ZK: First discovery of Cucubalus (Caryophyllaceae) fossil, and its biogeographical and ecological implications. Review of Palaeobotany and Palynology 2013, 190:41-47. 

88.       Zhang MG, Zhou ZK, Chen WY, Slik JWF, Cannon CH, Raes N: Using species distribution modeling to improve conservation and land use planning of Yunnan, China. Biological Conservation 2012, 153:257-264. 

89.       Yu WT, Jacques FMB, Chen ST, Zhou ZK: Nutlet micro-morphology of the genus Microula (Boraginaceae) from the Qinghai-Tibetan Plateau, and its systematic implications. Nordic Journal of Botany 2012, 30(5):596-612. 

90.       Xing YW, Utescher T, Jacques FMB, Su T, Liu Y, Huang YJ, Zhou ZK: Paleoclimatic estimation reveals a weak winter monsoon in southwestern China during the late Miocene: Evidence from plant macrofossils. Palaeogeography Palaeoclimatology Palaeoecology 2012, 358:19-26. 

91.       Xia K, Daws MI, Stuppy W, Zhou ZK, Pritchard HW: Rates of Water Loss and Uptake in Recalcitrant Fruits of Quercus Species Are Determined by Pericarp Anatomy. Plos One 2012, 7(10). 

92.       Xia K, Daws MI, Hay FR, Chen WY, Zhou ZK, Pritchard HW: A comparative study of desiccation responses of seeds of Asian Evergreen Oaks, Quercus subgenus Cyclobalanopsis and Quercus subgenus Quercus. South African Journal of Botany 2012, 78:47-54. 

93.       Pu CX, Chen WY, Zhou ZK: Elsholtzia litangensis sp nov (Lamiaceae) endemic to China. Nordic Journal of Botany 2012, 30(2):174-177. 

94.       Huang YJ, Jacques FMB, Liu YS, Su T, Xing YW, Xiao XH, Zhou ZK: New fossil endocarps of Sambucus (Adoxaceae) from the upper Pliocene in SW China. Review of Palaeobotany and Palynology 2012, 171:152-163. 

95.       Guo SX, Kvacek Z, Manchester SR, Zhou ZK: Ditaxocladus (extinct Cupressaceae, Cupressoideae) from the Upper Cretaceous and Paleocene of the Northern Hemisphere. Palaeontographica Abteilung B-Palaophytologie 2012, 288(5-6):135-159. 

96.       Chen ST, Xing YW, Su T, Zhou ZK, Dilcher EDL, Soltis DE: Phylogeographic analysis reveals significant spatial genetic structure of Incarvillea sinensis as a product of mountain building. Bmc Plant Biology 2012, 12. 

97.       Yu WT, Chen ST, Zhou ZK: Microula pentagona sp nov and M-galactantha sp nov (Boraginaceae) from the eastern Qinghai-Tibetan Plateau. Nordic Journal of Botany 2011, 29(2):215-220. 

98.       Yang QS, Te M, Gao JG, Zhou ZK: Original plant of Tibetan popular flower Skal-bzang Me-tog; 2011. 

99.       Su T, Jacques FMB, Liu YS, Xiang JY, Xing YW, Huang YJ, Zhou ZK: A new Drynaria (Polypodiaceae) from the Upper Pliocene of Southwest China. Review of Palaeobotany and Palynology 2011, 164(1-2):132-142. 

100.     Liu YS, Utescher T, Zhou ZK, Sun BN: The evolution of Miocene climates in North China: Preliminary results of quantitative reconstructions from plant fossil records. Palaeogeography Palaeoclimatology Palaeoecology 2011, 304(3-4):308-317. 

101.     Jacques FMB, Wang W, Ortiz RD, Li HL, Zhou ZK, Chen ZD: Integrating fossils in a molecular-based phylogeny and testing them as calibration points for divergence time estimates in Menispermaceae. Journal of Systematics and Evolution 2011, 49(1):25-49. 

102.     Jacques FMB, Su T, Spicer RA, Xing YW, Huang YJ, Wang WM, Zhou ZK: Leaf physiognomy and climate: Are monsoon systems different? Global and Planetary Change 2011, 76(1-2):56-62. 

103.     Jacques FMB, Liu CYS, Martinetto E, Zhou ZK: Revised taxonomy of selected fossil endocarp species in the Menispermaceae using a morphometric approach. Geodiversitas 2011, 33(1):177-197. 

104.     Jacques FMB, Guo SX, Su T, Xing YW, Huang YJ, Liu YS, Ferguson DK, Zhou ZK: Quantitative reconstruction of the Late Miocene monsoon climates of southwest China: A case study of the Lincang flora from Yunnan Province. Palaeogeography Palaeoclimatology Palaeoecology 2011, 304(3-4):318-327. 

105.     Yang Y, Zhou ZK: New insights into the species problem. Science China-Life Sciences 2010, 53(8):964-972. 

106.     Xing YW, Liu YS, Su T, Jacques FMB, Zhou ZK: Pinus prekesiya sp nov from the upper Miocene of Yunnan, southwestern China and its biogeographical implications. Review of Palaeobotany and Palynology 2010, 160(1-2):1-9. 

107.     Xia K, Seal CE, Chen WY, Zhou ZK, Pritchard HW: Fruit oil contents of the genus Quercus (Fagaceae): A comparative study on acorns of subgenus Quercus and the Asian subgenus Cyclobalanopsis. Seed Science and Technology 2010, 38(1):136-145. 

108.     Su T, Xing YW, Liu YS, Jacques FMB, Chen WY, Huang YJ, Zhou ZK: LEAF ''>Palaios 2010, 25(3-4):234-238. 

109.     Jacques FMB, Zhou ZK: Geometric morphometrics: A powerful tool for the study of shape evolution in Menispermaceae endocarps. Taxon 2010, 59(3):881-895. 

110.     Deng M, Zhou ZK, Coombes A: Lectotypification and New Synonymy in Quercus subg. Cyclobalanopsis (Fagaceae). Novon 2010, 20(4):400-405. 

111.      Chen ST, Gong J, Guan KY, Zhou ZK: Biodiversity Conservation of the Genus Incarvillea Juss. (Bignoniaceae) Based on Molecular Diversity and Species Richness Assessment. Journal of Plant Biology 2010, 53(6):387-394. 

112.     Ayodele AE, Zhou ZK: Scanning electron microscopy of fruits in the West African Polygonaceae. Journal of Systematics and Evolution 2010, 48(5):336-343. 

113.     Yang QS, Chen WY, Xia K, Zhou ZK: Climatic envelope of evergreen sclerophyllous oaks and their present distribution in the eastern Himalaya and Hengduan Mountains. Journal of Systematics and Evolution 2009, 47(3):183-190. 

114.     Xia K, Su T, Liu YS, Xing YW, Jacques FMB, Zhou ZK: Quantitative climate reconstructions of the late Miocene Xiaolongtan megaflora from Yunnan, southwest China. Palaeogeography Palaeoclimatology Palaeoecology 2009, 276(1-4):80-86. 

115.     Liu MQ, Deng M, Zhou ZK: Taxonomic and ecological implications of leaf cuticular morphology in Castanopsis, Castanea, and Chrysolepis. Plant Systematics and Evolution 2009, 283(1-2):111-123. 

116.     Li XX, Zhou ZK: Phylogenetic studies of the core Alismatales inferred from morphology and rbcL sequences. Progress in Natural Science-Materials International 2009, 19(8):931-945. 

117.     Zhang ZY, Du N, Pu CX, Gu ZJ, Zhou ZK: Environmental heterogeneity decides bio-heterogeneity of the Spiraea japonica complex (Rosaceae) in China. Australian Journal of Botany 2008, 56(1):91-95. 

118.     Xia K, Zhou ZK, Chen WY, Sun WB: Rescuing the Sichou oak Quercus sichourensis in China. Oryx 2008, 42(1):15-16. 

119.     Wang Y, Deng M, Zhang SY, Zhou ZK, Tian WX: Parasitic loranthus from Loranthaceae rather than Viscaceae potently inhibits fatty acid synthase and reduces body weight in mice. Journal of Ethnopharmacology 2008, 118(3):473-478. 

120.     Mitsui Y, Chen ST, Zhou ZK, Peng CI, Deng YF, Setoguchi H: Phylogeny and biogeography of the genus Ainsliaea (Asteraceae) in the Sino-Japanese region based on nuclear rDNA and plastid DNA sequence data. Annals of Botany 2008, 101(1):111-124. 

121.     Li HZ, Ma H, Zhou ZK, Guan KY: A new species of Begonia (Begoniaceae) from Guangxi, China. Botanical Journal of the Linnean Society 2008, 157(1):83-90. 


123.     Chen YQ, Deng M, Zhou ZK: A hypothesis on cupule evolution and the evidence from molecular phylogenies and fossils. Journal of Systematics and Evolution 2008, 46(1):41-52. 

124.     Zhou ZK, Yang QS, Xia K: Fossils of Quercus sect. Heterobalanus can help explain the uplift of the Himalayas. Chinese Science Bulletin 2007, 52(2):238-247. 

125.     Zhang SB, Zhou ZK, Hu H, Xu K: Gas exchange and resource utilization in two alpine oaks at different altitudes in the Hengduan Mountains. Canadian Journal of Forest Research-Revue Canadienne De Recherche Forestiere 2007, 37(7):1184-1193. 

126.     Ma CL, Moseley RK, Chen WY, Zhou ZK: Plant diversity and priority conservation areas of Northwestern Yunnan, China. Biodiversity and Conservation 2007, 16(3):757-774. 

127.     Li XX, Zhou ZK: The higher-level phylogeny of monocots based on matK, rbcL and 18S rDNA sequences. Acta Phytotaxonomica Sinica 2007, 45(2):113-133. 

128.     Li HM, Zhou ZK: Fossil nothofagaceous leaves from the Eocene of western Antarctica and their bearing on the origin, dispersal and systematics of Nothofagus. Science in China Series D-Earth Sciences 2007, 50(10):1525-1535. 

129.     Ji YH, Zhou ZK, Li H: Four new synonyms in the genus Paris (Trilliaceae). Acta Phytotaxonomica Sinica 2007, 45(3):388-390. 

130.     Zhou ZK, Yang XF, Yang QS: Land bridge and long-distance dispersal - Old views, new evidence. Chinese Science Bulletin 2006, 51(9):1030-1038. 

131.     Yang XF, Skidmore AK, Melick DR, Zhou ZK, Xu JC: Mapping non-wood forest product (matsutake mushrooms) using logistic regression and a GIS expert system. Ecological Modelling 2006, 198(1-2):208-218. 

132.     Yan N, Hu H, Huang JL, Xu K, Wang H, Zhou ZK: Micropropagation of Cypripedium flavum through multiple shoots of seedlings derived from mature seeds. Plant Cell Tissue and Organ Culture 2006, 84(1):113-117. 

133.     Uehara K, Tanaka N, Momohara A, Zhou ZK: Genetic diversity of an endangered aquatic plant, Potamogeton lucens subspecies sinicus. Aquatic Botany 2006, 85(4):350-354. 

134.    Manchester SR, Chen ZD, Zhou ZK: Wood anatomy of Craigia (Malvales) from southeastern Yunnan, China. Iawa Journal 2006, 27(2):129-136. 

135.     Ji YH, Zhou ZK, Li H: Paris xichouensis, a new combination of Trilliaceae from China. Acta Phytotaxonomica Sinica 2006, 44(5):612-613. 

136.     Ji YH, Li H, Zhou ZK: Paris caobangensis Y. H. Ji, H. Li & Z. K. Zhou (Trilliaceae), a new species from northern Vietnam. Acta Phytotaxonomica Sinica 2006, 44(6):700-703. 

137.     Ji YH, Fritsch PW, Li H, Xiao TJ, Zhou ZK: Phylogeny and classification of Paris (Melanthiaceae) inferred from DNA sequence data. Annals of Botany 2006, 98(1):245-256. 

138.     Deng M, Zhou ZK, Coombes A: Taxonomic notes on the genus Cyclobalanopsis (Fagaceae). Annales Botanici Fennici 2006, 43(1):57-61. 

139.     Chen ST, Guan KY, Zhou ZK: A new subgenus of Incarvillea (Bignoniaceae). Annales Botanici Fennici 2006, 43(4):288-290. 

140.     Zhang SB, Zhou ZK, Hu H, Xu K, Yan N, Li SY: Photosynthetic performances of Quercus pannosa vary with altitude in the Hengduan Mountains, southwest China. Forest Ecology and Management 2005, 212(1-3):291-301. 

141.     Zhang SB, Hu H, Zhou ZK, Xu K, Yan N, Li SY: Photosynthesis in relation to reproductive success of Cyptipedium flavum. Annals of Botany 2005, 96(1):43-49. 

142.     Zhang SB, Hu H, Zhou ZK, Xu K, Yan N, Li SY: Photosynthetic performances of transplanted Cypripedium flavum plants. Botanical Bulletin of Academia Sinica 2005, 46(4):307-313. 

143.     Li XX, Zhou ZK: Endemic wild ornamental plants from northwestern Yunnan, China. Hortscience 2005, 40(6):1612-1619. 

144.     Chen ST, Guan KY, Zhou ZK, Olmstead R, Cronk Q: Molecular phylogeny of Incarvillea (Bignoniaceae) based on its and trnL-F sequences. American Journal of Botany 2005, 92(4):625-633. 

145.     Chen ST, Zhou ZK, Guan KY, Nakata M: Karyomorphology of Incarvillea (Bignoniaceae) and its implications in distribution and taxonomy. Botanical Journal of the Linnean Society 2004, 144(1):113-121. 

146.     Zhang ZY, Zhou ZK, Gu ZJ: Karyomorphology of Heptacodium (Caprifoliaceae s.str.) and its phylogenetic implications. Taxon 2002, 51(3):499-505. 

147.     Shui YM, Zhang GF, Zhou ZK, Mo MZ: Sladenia integrifolia (Sladeniaceae), a new species from China. Novon 2002, 12(4):539-542. 

148.     Luo Y, Zhou ZK: Leaf architecture in Quercus subgenus Cyclobalanopsis (Fagaceae) from China. Botanical Journal of the Linnean Society 2002, 140(3):283-295. 

149.     Zhou ZK, Crepet WL, Nixon KC: The earliest fossil evidence of the Hamamelidaceae: Late Cretaceous (Turonian) inflorescences and fruits of Altingioideae. American Journal of Botany 2001, 88(5):753-766. 

150.     Manos PS, Zhou ZK, Cannon CH: Systematics of Fagaceae: Phylogenetic tests of reproductive trait evolution. International Journal of Plant Sciences 2001, 162(6):1361-1379. 

151.     Huang SH, Sun H, Zhou ZK: Agapetes subsessilifolia (Ericaceae), a new species from the eastern Himalayas. Novon 1998, 8(4):378-379. 

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